Inadvertent alterations of the earth's atmosphere by man's activities are now of regional and even global proportion. Increasing concern has been focused in the last decade on consequences of acid rain, carbon dioxide enrichment of the atmosphere and reduction of ozone in the upper atmosphere. The latter two problems are of truly global scale. This book focuses on the atmospheric ozone reduction problem and the potential consequences for plant life. unlike carbon dioxide enrichment, reduction of the total atmospheric ozone column has not yet taken place to a noticeable degree -- it is a problem of the future. The processes leading to ozone reduction involve time periods on the scale of decades. However, by the same token, if society finds ozone reduction to be unacceptable it will take even longer for the process to be reversed. Thus, anticipation of the consequences of ozone reduction is of obvious importance. Speculation of the possibility of ozone reduction first appeared in the early 1970's and was focused on the consequences of the injection of large quantities of nitrogen oxides into the upper atmosphere by supersonic aircraft flying at high altitudes. Other sources of nitrogen oxides originating from the earth's surface were also considered. With further refinement, the concerns of nitrogen oxide pollution of the upper atmosphere were diminished since the quantities likely to be involved were insufficient to cause a serious threat to the ozone layer.

Current phase-out schedules of the production and emission of CFC's indicate that chlorine loading in the stratosphere is not yet at its maximum. The recovery of stratospheric ozone is estimated to take time and ele vated levels of UV-B radiation are expected to occur throughout most of the next century. Despite numerous physiological studies of UV-B effects on plants, often grown in climate chambers, knowledge of UV-B effects on organisms and processes in natural aquatic or terrestrial ecosystems is poor. Currently it appears that UV-B radiation is not just an environmental stress' factor to plants. In various ways, which are incompletely understood, UV-B affects a wide range of physiological and ecological processes. Remarkably, recent field studies indicate that enhanced UV-B does not markedly affect photosynthesis, growth and primary production, but rather interferes with plant morphogenesis and plant and ecosystem functions relating to the secondary metabolism. This special issue and book UV-B and Biosphere is an attempt to cover this range and to report the progress made in the research of ecological effects of enhanced solar UV-B radiation. The papers in this book formed the basis of an international workshop entitled' UV-B and Biosphere' , December 15-18, 1995, in Wageningen, The Netherlands. A first reaction of Hans de Boois on the number of papers and sessions scheduled from Friday to Sunday morning was: far too many.

From microbial to plant ecosystems, this book examines how changes in UV radiation, caused by anthropogenic ozone depletion, as well as changes in radiation levels throughout the evolution of life on Earth, can alter species composition and interspecies competitiveness. It focuses on the evolutionary aspects of the effects of UV as well as the various synergistic interactions of UV radiation with other environmental factors. Because our knowledge of UV effects on whole ecosystems is still at a relatively early stage, an important part of each chapter is an overview of future research directions and indications of where new date and knowledge is needed.

The research in this book covers papers on a great number of research projects on the responses of plants and crops of natural terrestrial ecosystems, of agro-ecosystems, and of aquatic ecosystems, to enhanced solar UV-B as a result of stratospheric ozone depletion. Some introductory chapters deal with general aspects of how plants respond to UV-B radiation. Photosynthetically Active Radiation (PAR) is a primary energy resource for terrestrial plants, necessary for plant growth. Inevitably terrestrial plants absorb UV-B when exposed to solar radiation. The spectral balance between PAR and UV-B is discussed in several chapters. The responses of plants and ecosystems from the Antarctic and Arctic to enhanced solar UV-B radiation as a consequence of the hole in the ozone layer are considered in some detail. In addition the papers in the book discuss the problem of how responses of plants to UV-B radiation interact with other environmental factors. The book is of great importance for those who are involved in global change topics: biologists, ecologists, earth scientists, agronomists, environmental scientists, and those who develop environmental policy.