Recent advances in statistical approaches called phylogenetic comparative methods (PCMs) have provided paleontologists with a powerful set of analytical tools for investigating evolutionary tempo and mode in fossil lineages. However, attempts to integrate PCMs with fossil data often present workers with practical challenges or unfamiliar literature. This Element presents guides to the theory behind and the application of PCMs with fossil taxa. Based on an empirical dataset of Paleozoic crinoids, example analyses are presented to illustrate common applications of PCMs to fossil data, including investigating patterns of correlated trait evolution and macroevolutionary models of morphological change. The authors emphasize the importance of accounting for sources of uncertainty and discuss how to evaluate model fit and adequacy. Finally, the authors discuss several promising methods for modeling heterogeneous evolutionary dynamics with fossil phylogenies. Integrating phylogeny-based approaches with the fossil record provides a rigorous, quantitative perspective on understanding key patterns in the history of life.

An authoritative introduction to the latest comparative methods in evolutionary biology Phylogenetic comparative methods are a suite of statistical approaches that enable biologists to analyze and better understand the evolutionary tree of life, and shed vital new light on patterns of divergence and common ancestry among all species on Earth. This textbook shows how to carry out phylogenetic comparative analyses in the R statistical computing environment. Liam Revell and Luke Harmon provide an incisive conceptual overview of each method along with worked examples using real data and challenge problems that encourage students to learn by doing. By working through this book, students will gain a solid foundation in these methods and develop the skills they need to interpret patterns in the tree of life. Covers every major method of modern phylogenetic comparative analysis in R Explains the basics of R and discusses topics such as trait evolution, diversification, trait-dependent diversification, biogeography, and visualization Features a wealth of exercises and challenge problems Serves as an invaluable resource for students and researchers, with applications in ecology, evolution, anthropology, disease transmission, conservation biology, and a host of other areas Written by two of today’s leading developers of phylogenetic comparative methods

All-new edition of the world’s leading vertebrate palaeontology textbook, now addressing key evolutionary transitions and ecological drivers for vertebrate evolution Richly illustrated with colour illustrations of the key species and cladograms of all major vertebrate taxa, Vertebrate Palaeontology provides a complete account of the evolution of vertebrates, including macroevolutionary trends and drivers that have shaped their organs and body plans, key transitions such as terrestrialization, endothermy, flight and impacts of mass extinctions on biodiversity and ecological drivers behind the origin of chordates and vertebrates, their limbs, jaws, feathers, and hairs. This revised and updated fifth edition features numerous recent examples of breakthrough discoveries in line with the current macroevolutionary approach in palaeontology research, such as the evolutionary drivers that have shaped vertebrate development. Didactical features have been enhanced and include new functional and developmental feature spreads, key questions, and extensive references to useful websites. Written by a leading academic in the field, Vertebrate Palaeontology discusses topics such as: Palaeozoic fishes, including Cambrian vertebrates, placoderms (‘armour-plated monsters’), Pan-Chondrichthyes such as sharks and rays, and Osteichthyes (‘bony fishes’) The first tetrapods, covering problems of life on land, diversity of Carboniferous tetrapods and temnospondyls and reptiliomorphs following the Carboniferous Mesozoic reptiles, such as Testudinata (turtles), Crocodylomorpha, Pterosauria, Dinosauria, great sea dragons and Lepidosauria (lizards and snakes) Mammals of the southern and northern hemispheres, covering Xenarthra (sloths, anteaters), Afrotheria (African mammals), Laurasiatheria (bats, ungulates, carnivores), and Euarchontoglires (rodents, primates) A highly comprehensive and completely up-to-date reference on vertebrate evolution, Vertebrate Palaeontology is an ideal learning aid for palaeontology courses in biology and geology departments. The text is also highly valuable to enthusiasts who want to experience the flavour of how modern research in the field is conducted.

Macroevolutionary inference has historically been treated as a two-step process, involving the inference of a tree, and then inference of a macroevolutionary model using that tree. Newer models blend the two steps. These methods make more complete use of fossils than the previous generation of Bayesian phylogenetic models. They also involve many more parameters than prior models, including parameters about which empiricists may have little intuition. In this Element, we set forth a framework for fitting complex, hierarchical models. The authors ultimately fit and use a joint tree and diversification model to estimate a dated phylogeny of the Cincta (Echinodermata), a morphologically distinct group of Cambrian echinoderms that lack the fivefold radial symmetry characteristic of extant members of the phylum. Although the phylogeny of cinctans remains poorly supported in places, this Element shows how models of character change and diversification contribute to understanding patterns of phylogenetic relatedness and testing macroevolutionary hypotheses.

Fossil crinoids are exceptionally suited to deep-time studies of community paleoecology and niche partitioning. By merging ecomorphological trait and phylogenetic data, this Element summarizes niche occupation and community paleoecology of crinoids from the Bromide fauna of Oklahoma (Sandbian, Upper Ordovician). Patterns of community structure and niche evolution are evaluated over a ~5 million-year period through comparison with the Brechin Lagerstätte (Katian, Upper Ordovician). The authors establish filtration fan density, food size selectivity, and body size as major axes defining niche differentiation, and niche occupation is strongly controlled by phylogeny. Ecological strategies were relatively static over the study interval at high taxonomic scales, but niche differentiation and specialization increased in most subclades. Changes in disparity and species richness indicate the transition between the early-middle Paleozoic Crinoid Evolutionary Faunas was already underway by the Katian due to ecological drivers and was not triggered by the Late Ordovician mass extinction.

Phylogenetic analysis and morphometrics have been developed by biologists into rigorous analytic tools for testing hypotheses about the relationships between groups of species. This book applies these tools to paleontological data. The fossil record is our one true chronicle of the history of life, preserving a set of macroevolutionary patterns; thus various hypotheses about evolutionary processes can be tested in the fossil record using phylogentic analysis and morphometrics. The first book of its type, Fossils, Phylogeny, and Form will be useful in evolutionary biology, paleontology, systematics, evolutionary development, theoretical biology, biogeography, and zoology. It will also provide a practical, researcher-friendly gateway into computer-based phylogenetics and morphometrics.

This Element reviews the ecologies of skeletal trace-producing interactions on echinoids in Modern ecosystems and the recognition of those biogenic traces in the fossil record. This title is also available as Open Access on Cambridge Core.